[Taxacom] Dipsacales classification (Subclass Asteridae)

Kenneth Kinman kennethkinman at webtv.net
Mon Apr 5 21:09:12 CDT 2010


 Dear All,       
      Having preliminarily examined what gene sequences exist for the
purported "clade"  Paracryphiales (which are not many), the present
paucity of data makes it unclear if it is really a clade or if it is
paraphyletic.  In any case, it seems likely that Quintiniaceae (along
with it purported fossil relative Silvianthemum) are the basalmost
members of Dipsacales sensu lato.           
       For the time being (until more information becomes available), I
am coding Plesion Silvianthemum as the basalmost clade, followed by an
extant clade (equivalent to APG's Paracryphiales) with Quintiniaceae at
its base, and then what I call the "core Dipsacales".  Of course, it is
also possible that Silvianthemum clades exclusively with Quintiniaceae,
and then a Paracryphiaceae-Sphenostemonaceae clade  splitting off next,
but this may be less likely from a biogeographic perspective.  Family
Quintiniaceae most likely originated in what is now mainland Asia, with
the modern relicts only surviving offshore on SW Pacific islands (along
with relatives in Sphenostemonaceae and Paracryphiaceae).  The earliest
"core Dipsacales" also presumably originated in Asia (or Europe) as
well.

      But there simply doesn't seem to be enough data to precisely code
their exact phylogenetic relationships with any certainty at the present
time.  However, I still feel that this slight expansion of Order
Dipsacales (as a sister group to Order Apiales) is phylogenetically the
best option against which to evaluate and incorporate additional data.


Dipsacales 
    1  Plesion Silvianthemum (Cretaceous fossil)
    2  Quintiniaceae 
    B  Sphenostemonaceae
    C  Paracryphiaceae   
    3  Viburnaceae 
    B  Sambucaceae 
    C  Adoxaceae 
    4  Diervillaceae 
    5  Caprifoliaceae 
    6  Linnaeaceae 
    7  Morinaceae 
    8  Dipsacaceae 
    9  Valerianaceae   

                  --------Ken Kinman 





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